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  • UC Riverside
  • College of Natural and Agricultural Sciences

Biology Department, University of California, Riverside

Complete list of technical publications, Helen M. Regan (helen.regan@ucr.edu )


Journal articles (* denotes student authorship)

  1. Regan, H.M., D.A. Keith, T.J. Regan, M.G. Tozer & N. Tootell*. Fire management to combat disease: turning interactions between threats into conservation management. (Accepted Oecologia)
  2. Syphard, A.D., K.C. Clarke, J. Franklin, H.M. Regan, and M. McGinnis. Forecasts of Habitat Loss and Fragmentation Due to Urban Growth are Sensitive to Input Data Quality and Scale. (In press Journal of Environmental Management)
  3. Martínez-Abraín, A., H.M. Regan, C. Viedma, E. Villuendas, M.A. Bartolomé, J. A. Gómez and D. Oro. Cost-effectiveness of translocation options for a threatened waterbird. (In press Conservation Biology)
  4. Mizerek, T.*, H.M. Regan, and K.A. Hovel. Seagrass habitat loss and fragmentation influence optimal management strategies for a blue crab (Callinectes sapidus) fishery. Marine Ecology Progress Series 427: 247–257, 2011.
  5. Franklin, J., H.M. Regan, L.A. Hierl, D.H. Deutschman, B.S. Johnson, and C.S. Winchell. Planning, implementation and monitoring of multiple species habitat conservation plans. American Journal of Botany 98:559-571, 2011.
  6. Colyvan, M., J. Justus, and H.M. Regan. The conservation game. Biological Conservation 144(4):1246-1253, 2011.
  7. Colyvan, M., J. Justus, and H.M. Regan. The natural environment is valuable but not infinitely valuable. Conservation Letters 3(4):224-228, 2010.
  8. Foley, M.M., B.S. Halpern, F. Micheli, M.H. Armsby, M.R. Caldwell, C.M. Crain, E. Prahler, D. Sivas, N. Rohr, M.W. Beck, M.H. Carr, L.B. Crowder, J.E. Duffy, S.D. Hacker, K. McLeod, C.H. Peterson, H.M. Regan, M.H. Ruckelshaus, P.A. Sandifer, R.S. Steneck. Guiding ecological principles for marine spatial planning. Marine Policy 35(5):955-966, 2010.
  9. Lawson, D. M.*, H.M. Regan, P.H. Zedler and J. Franklin. Cumulative effects of land use, altered fire regime and climate change on persistence of Ceanothus verrucosus, a rare, fire-dependent plant species. Global Change Biology 16(9):2518–2529, 2010.
  10. Ellstrand, N.C., D. Biggs, A. Kaus, P. Lubinsky, L.A. McDade, K. Preston, L. Prince, H.M. Regan, V. Rorive, O. Ryder. Got Hybridization? A Multidisciplinary Approach for Informing Science Policy. BioScience 60:384-388, 2010.
  11. Regan, H.M., J.B. Crookston*, R. Swab*, J. Franklin, and D.M. Lawson*. Habitat fragmentation and altered fire regime create trade-offs for the persistence of an obligate seeding shrub. Ecology 91(4):1114-1123, 2010.
  12. Justus, J., M. Colyvan, H.M. Regan and L.A. Maguire. Response to Sagoff. Trends in Ecology and Evolution 24(12):644, 2009.
  13. Justus, J., M. Colyvan, H.M. Regan and L.A. Maguire. Buying into conservation: intrinsic versus instrumental value. Trends in Ecology and Evolution 24(4):187-191, 2009.
  14. Keith, D.A., H.R. Akçakaya, W. Thuiller, G.F. Midgley, R.G. Pearson, S.J. Phillips, H.M. Regan, M.B. Araújo, T.G. Rebelo. Predicting extinction risks under climate change: a new mechanistic approach linking stochastic population models with dynamic bioclimatic habitat models. Biology Letters 4:560-563, 2008.
  15. Hierl, L.A., J. Franklin, D.H. Deutschman, H.M. Regan, and B.S. Johnson. Assessing and prioritizing ecological communities for monitoring in a regional habitat conservation plan. Environmental Management 42(1):165–179, 2008.
  16. Regan, H.M., L.A. Hierl, J. Franklin, D.H. Deutschman, H.L. Schmalbach, C.S. Winchell and B.S. Johnson. Species prioritisation for monitoring and management in regional multiple species conservation plans. Diversity and Distributions, 14(4): 262-271, 2008.
  17. Hovel, K.A. and H.M. Regan. Using an individual-based model to examine the roles of habitat fragmentation and behavior on predator-prey relationships in seagrass landscapes. Landscape Ecology, 23(1):75-89, 2008.
  18. Markovchick-Nicholls*, L., H.M. Regan, D.H. Deutschman, A Widyanata*, B. Martin, L. Noreke and T.A. Hunt. Relationships between human disturbance and wildlife land use in urban habitat fragments. Conservation Biology, 22(1):99-109, 2008.
  19. Steele, K.S.*, H.M. Regan, M. Colyvan and M.A. Burgman. Right decisions or happy decision makers? Social Epistemology, 21(4):349-368, 2007.
  20. Colyvan, M. and H.M. Regan. Legal decisions and the reference class problem. International Journal of Evidence and Proof, 11(4):274–285, 2007. (invited article)
  21. Regan, H.M., F.W. Davis, S.J. Andelman, A. Widyanata* and M. Freese*. Comprehensive criteria for biodiversity evaluation in conservation planning. Biodiversity and Conservation 16:2715-2728, 2007.
  22. Keith, D.A., M.G. Tozer, T.J. Regan and H.M. Regan. The persistence niche: what makes it and what breaks it for two fire-prone plant species from south-eastern Australia. Advances in Plant Conservation Special Issue. Australian Journal of Botany 55(3):273-279, 2007.
  23. Regan, H.M., M. Colyvan and L. Markovchick-Nicholls*. A formal model for consensus and negotiation in environmental management. Journal of Environmental Management 80:167-176, 2006.
  24. Halpern B.S., H.M. Regan, H.P. Possingham, M.A. McCarthy. Accounting for uncertainty in marine reserve design. Ecology Letters 9:2-11, 2006. (invited article)
  25. Halpern B.S., H.M. Regan, H.P. Possingham, M.A. McCarthy. Rejoinder: uncertainty and decision making. Ecology Letters 9(1):13-14, 2006. (invited article)
  26. Regan, H.M., Y. Ben-Haim, B. Langford, W.G. Wilson, P. Lundberg, S.J. Andelman, and M.A. Burgman. Robust decision making under severe uncertainty for conservation management. Ecological Applications 15(4):1471-1477, 2005.
  27. Keith D.A., M.A. McCarthy, H.M. Regan, T.J. Regan*, C. Bowles*, C. Drill, C. Craig*, B. Pellow, M.A. Burgman, L.L. Master, M. Ruckelshaus, B. McKenzie, S.J. Andelman, P.R. Wade. Protocols for listing threatened species can forecast extinction. Ecology Letters 7:1101-1108, 2004.
  28. Andelman, S.J., C. Groves, and H.M. Regan. A review of the U.S. Forest Service's selection process of species at risk for viability assessments. Acta Oecologica, 26:75-83, 2004.
  29. McCarthy, M.A., D.A. Keith, J. Tietjen*, M.A. Burgman, M. Maunder, L. Master, B.W. Brook, G. Mace, H.P. Possingham, R. Medellin, S.J. Andelman, H.M. Regan, T.J. Regan*, and M. Ruckleshaus. Comparing predictions of extinction risk using models and subjective judgement. Acta Oecologica, 26:67-74, 2004.
  30. Regan, H.M., S. Ferson and D. Berleant. Equivalence of methods for uncertainty propagation of real-valued random variables. International Journal of Approximate Reasoning, 36(1):1-30, 2004.
  31. Pastorok, R.A., H.R. Akçakaya, H.M. Regan, S. Ferson, and S.M. Bartell. Role of ecological modeling in risk assessment. Human and Ecological Risk Assessment, 9(4):939-972, 2003.
  32. Bartell, S.M., R.A. Pastorok, H.R. Akçakaya, H.M. Regan, S. Ferson and C. Mackay. Realism and relevance of ecological models used in chemical risk assessment. Human and Ecological Risk Assessment, 9(4):907-938, 2003.
  33. Regan, H.M., H.R. Akçakaya, S. Ferson, K.V. Root, S. Carroll* and L.R. Ginzburg. Treatments of uncertainty and variability in ecological risk assessment of single-species populations. Human and Ecological Risk Assessment, 9(4):889-906, 2003.
  34. Taylor, R.J., T.J. Regan*, H.M. Regan, M.A. Burgman and K.Bonham*. Impacts of plantation development, harvesting schedules and rotation lengths on the rare snail Tasmaphena lamproides in northwest Tasmania: a population viability analysis. Forest Ecology and Management 175:455-466, 2003.
  35. Regan, H.M., T.D. Auld, D. Keith and M.A. Burgman. The effects of fire and predators on the long-term persistence of an endangered shrub Grevillea caleyi. Biological Conservation 109(1):73-83, 2003.
  36. Regan, H.M., B.K. Hope, and S. Ferson. Analysis and portrayal of uncertainty in a food web exposure model. Human and Ecological Risk Assessment, 8(7):1757-1777, 2002. Awarded Ecological Risk Assessment Paper of Year 2002 in Journal of Human and Ecological Risk Assessment
  37. Regan, H.M. Von Neumann stability analysis of symplectic integrators applied to Hamiltonian PDEs. Journal of Computational Mathematics, 20(6):611-618, 2002.
  38. Elith, J.*, M.A. Burgman and H.M. Regan. Mapping epistemic uncertainty and vague concepts in predictions of species' distribution. Ecological Modelling, 157:313-329, 2002.
  39. Regan, H.M., B.E. Sample, and S. Ferson. Deterministic and Probabilistic Ecological Soil Screening Levels for Wildlife. Environmental Toxicology and Chemistry, 21(4):882-890, 2002.
  40. Regan, H.M., M. Colyvan, and M.A. Burgman. A taxonomy and treatment of uncertainty for ecology and conservation biology. Ecological Applications, 12(2):618-628, 2002.
  41. Colyvan, M., H.M. Regan, and S. Ferson. Is it a crime to belong to a reference class?, The Journal of Political Philosophy, 9(2):168-181, 2001.
  42. Regan, T.J.*, H.M. Regan, K. Bonham*, R.J. Taylor, and M.A. Burgman. Modelling the impact of timber harvesting on a rare carnivorous land snail (Tasmaphena lamproides) in northwest Tasmania, Australia. Ecological Modelling, 139:253-264, 2001.
  43. Regan, H.M., R. Lupia, A.N. Drinnan and M.A. Burgman. The currency and tempo of extinction. The American Naturalist, 157(1):1-10, January 2001. Highlighted in Scientific American, 285(5):40-49 Nov 2001 and The Age, 10th Oct 1998.
  44. Regan, H.M., and M. Colyvan. Fuzzy Sets and Threatened Species Classification, Conservation Biology, 14(4):1197-1199, August 2000.
  45. Regan, H.M., M. Colyvan and M.A. Burgman. A Proposal for Fuzzy IUCN Categories and Criteria, Biological Conservation, 92(1):101-108, 2000.
  46. Bearlin, A.R.*, M.A. Burgman, and H.M. Regan. A Stochastic Model for Seagrass (Zostera muelleri) in Port Phillip Bay, Victoria, Australia, Ecological Modelling, 118:131-148, 1999.
  47. Stiles, P.J. and H.M. Regan. Transient Cellular Convection in Electrically Polarized Colloidal Suspensions, Journal of Colloid and Interface Science, 202(2):562-565, 1998.


Book Chapters

  1. Regan, H.M., M. Ensbey* and M.A. Burgman. Chapter 11: Conservation prioritisation and uncertainty in planning inputs. In A. Moilanen, K.A. Wilson and H.P. Possingham (eds.) Spatial Conservation Prioritisation: Quantitative Methods and Computational Tools. Oxford University Press. Pp. 145-157, 2009.
  2. Hayes, K., H.M. Regan and M.A. Burgman. Introduction to the concepts and methods of uncertainty analysis. In A.R. Kapuscinski, K.R. Hayes, L. Si, and G. Dana (eds.) Environmental Risk Assessment of Genetically Modified Organisms, Volume 3: Methodologies for Transgenic Fish. CAB International, Wallingford, UK. pp. 188-208, 2007.
  3. Gervais, J. and H.M. Regan. What conservation biology and natural resource management can offer population-level ecological risk assessment. In: Barnthouse LW, Munns WRM, and Sorensen MT (editors). Population-Level Ecological Risk Assessment. SETAC Press, Pensacola, FL, USA, pp.129-150, 2007.
  4. Menzie, C., N. Bettinger, A. Fritz, L. Kapustka, H.M. Regan, V. Møller, H. Noel*. Population protection goals. In: Barnthouse LW, Munns WRM, and Sorensen MT (editors). Population-Level Ecological Risk Assessment. SETAC Press, Pensacola, FL, USA. pp. 41-68, 2007.
  5. Regan, H.M. and T.D. Auld. Using Population Viability Analysis for Management of an Endangered Australian Shrub, Grevillea caleyi. In H.R. Akçakaya, M.A. Burgman, O. Kindvall, C.C. Wood, P. Sjogren-Gulve, J. Hatfield, and M. McCarthy (eds.), Species Conservation and Management: Case Studies, Oxford University Press, pp. 23-35, 2004.
  6. Colyvan, M., H.M. Regan, and S. Ferson. Is it a crime to belong to a reference class?, In H. Kyburg and M. Thalos (eds.) Probability is the Very Guide in Life, Open Court, Chicago, pp. 331-347, 2003. (reprinted from journal article)
  7. Regan, H.M. Population Models: Individual-Based, in R.A. Pastorok, S.M. Bartell, S. Ferson, L.R. Ginzburg (eds.) Ecological Modeling in Risk Assessment: Chemical Effects on Populations, Ecosystems and Landscapes, Lewis Publishers, Boca Raton FL., pp. 65-82, 2002.
  8. Akcakaya, H.R. and H.M. Regan. Population Models: Metapopulations, in R.A. Pastorok, S.M. Bartell, S. Ferson, L.R. Ginzburg (eds.) Ecological Modeling in Risk Assessment: Chemical Effects on Populations, Ecosystems and Landscapes, Lewis Publishers, Boca Raton FL., pp. 83-95, 2002.